Introduction

During the SEDAR 46 stock assessment for US Caribbean Data-limited Species (2015-2016), candidate management procedures were evaluated and tested (via management strategy evaluation) for their feasibility in providing catch advice for six stocks.

This operating model was based on parameters and data reported in Sagarese et al. (2018) and SEDAR (2016).

Figure 1. The U.S. Carribean EEZ and Council boundaries (image taken from SEDAR 2016)

Figure 2. Queen triggerfish (image courtesy of the NOAA Photo Library)

Operating Model

The OM rdata file can be downloaded from here

Download and import into R using myOM <- readRDS('OM.rdata')

Species Information

Species: Balistes vetula

Common Name: Queen Triggerfish

Management Agency: NOAA

Region: U.S. Caribbean

Sponsor: NOAA

Latitude: 18

Longitude: -65

OM Parameters

OM Name: Name of the operating model: Queen Triggerfish

nsim: The number of simulations: 192

proyears: The number of projected years: 40

interval: The assessment interval - how often would you like to update the management system? 3

pstar: The percentile of the sample of the management recommendation for each method: 0.5

maxF: Maximum instantaneous fishing mortality rate that may be simulated for any given age class: 3

reps: Number of samples of the management recommendation for each method. Note that when this is set to 1, the mean value of the data inputs is used. 1

Source: A reference to a website or article from which parameters were taken to define the operating model

Sagarese et al. (2018) N. Amer. J. Fish. Manage. 38:507-537

Stock Parameters

Mortality and age: maxage, R0, M, M2, Mexp, Msd

maxage: The maximum age of individuals that is simulated (there is no plus group ). Single value. Positive integer

Specified Value(s): 14

Based on Sagarese et al. (2018).

R0: The magnitude of unfished recruitment. Single value. Positive real number

Specified Value(s): 1000

Arbitrary value chosen for scaling population size.

M: Natural mortality rate. Uniform distribution lower and upper bounds. Positive real number

Specified Value(s): 0.3, 0.47

Based on Sagarese et al. (2018).

M2: (Optional) Natural mortality rate at age. Vector of length maxage . Positive real number

Slot not used.

Mexp: Exponent of the Lorenzen function assuming an inverse relationship between M and weight. Uniform distribution lower and upper bounds. Real numbers <= 0.

Specified Value(s): 0, 0

Slot not used. Age-constant M was assumed.

Msd: Inter-annual variability in natural mortality rate expressed as a coefficient of variation. Uniform distribution lower and upper bounds. Non-negative real numbers

Specified Value(s): 0, 0.05

From Sagarese et al. (2018).

Natural Mortality Parameters

Sampled Parameters

Histograms of 48 simulations of M, Mexp, and Msd parameters, with vertical colored lines indicating 3 randomly drawn values used in other plots:

Time-Series

The average natural mortality rate by year for adult fish for 3 simulations. The vertical dashed line indicates the end of the historical period:

M-at-Age

Natural mortality-at-age for 3 simulations in the first historical year, the last historical year (i.e., current year), and the last projected year:

M-at-Length

Natural mortality-at-length for 3 simulations in the first historical year, the last historical year (i.e., current year), and the last projected year:

Recruitment: h, SRrel, Perr, AC

h: Steepness of the stock recruit relationship. Uniform distribution lower and upper bounds. Values from 1/5 to 1

Specified Value(s): 0.35, 0.84

From Sagarese et al. (2018).

SRrel: Type of stock-recruit relationship. Single value, switch (1) Beverton-Holt (2) Ricker. Integer

Specified Value(s): 1

From Sagarese et al. (2018).

Perr: Process error, the CV of lognormal recruitment deviations. Uniform distribution lower and upper bounds. Non-negative real numbers

Specified Value(s): 0.2, 0.5

From Sagarese et al. (2018).

AC: Autocorrelation in recruitment deviations rec(t)=ACrec(t-1)+(1-AC)sigma(t). Uniform distribution lower and upper bounds. Non-negative real numbers

Specified Value(s): 0.1, 0.9

From Sagarese et al. (2018).

Recruitment Parameters

Sampled Parameters

Histograms of 48 simulations of steepness (h), recruitment process error (Perr) and auto-correlation (AC) for the Beverton-Holt stock-recruitment relationship, with vertical colored lines indicating 3 randomly drawn values used in other plots:

Time-Series

Non-stationarity in stock productivity: Period, Amplitude

Period: (Optional) Period for cyclical recruitment pattern in years. Uniform distribution lower and upper bounds. Non-negative real numbers

Slot not used.

Amplitude: (Optional) Amplitude in deviation from long-term average recruitment during recruitment cycle (eg a range from 0 to 1 means recruitment decreases or increases by up to 100% each cycle). Uniform distribution lower and upper bounds. 0 < Amplitude < 1

Slot not used.

Growth: Linf, K, t0, LenCV, Ksd, Linfsd

Linf: Maximum length. Uniform distribution lower and upper bounds. Positive real numbers

Specified Value(s): 415, 605

From Sagarese et al. (2018).

K: von Bertalanffy growth parameter k. Uniform distribution lower and upper bounds. Positive real numbers

Specified Value(s): 0.14, 0.4

From Sagarese et al. (2018).

t0: von Bertalanffy theoretical age at length zero. Uniform distribution lower and upper bounds. Non-positive real numbers

Specified Value(s): -1.8, -0.6

From Sagarese et al. (2018).

LenCV: Coefficient of variation of length-at-age (assumed constant for all age classes). Uniform distribution lower and upper bounds. Positive real numbers

Specified Value(s): 0.13, 0.25

From Sagarese et al. (2018).

Ksd: Inter-annual variability in growth parameter k expressed as coefficient of variation. Uniform distribution lower and upper bounds. Non-negative real numbers

Specified Value(s): 0, 0.02

From Sagarese et al. (2018).

Linfsd: Inter-annual variability in maximum length expressed as a coefficient of variation. Uniform distribution lower and upper bounds. Non-negative real numbers

Specified Value(s): 0, 0

Slot not used.

Growth Parameters

Sampled Parameters

Histograms of 48 simulations of von Bertalanffy growth parameters Linf, K, and t0, and inter-annual variability in Linf and K (Linfsd and Ksd), with vertical colored lines indicating 3 randomly drawn values used in other plots:

Time-Series

The Linf and K parameters in each year for 3 simulations. The vertical dashed line indicates the end of the historical period:

Growth Curves

Sampled length-at-age curves for 3 simulations in the first historical year, the last historical year, and the last projection year.

Maturity: L50, L50_95

L50: Length at 50 percent maturity. Uniform distribution lower and upper bounds. Positive real numbers

Specified Value(s): 215, 235

From Sagarese et al. (2018).

L50_95: Length increment from 50 percent to 95 percent maturity. Uniform distribution lower and upper bounds. Positive real numbers

Specified Value(s): 45, 65

From Sagarese et al. (2018).

Maturity Parameters

Sampled Parameters

Histograms of 48 simulations of L50 (length at 50% maturity), L95 (length at 95% maturity), and corresponding derived age at maturity parameters (A50 and A95), with vertical colored lines indicating 3 randomly drawn values used in other plots:

Maturity at Age and Length

Maturity-at-age and -length for 3 simulations in the first historical year, the last historical year (i.e., current year), and the last projected year:

Stock depletion and Discard Mortality: D, Fdisc

D: Current level of stock depletion SSB(current)/SSB(unfished). Uniform distribution lower and upper bounds. Fraction

Specified Value(s): 0.1, 0.54

From Sagarese et al. (2018).

Fdisc: Fraction of discarded fish that die. Uniform distribution lower and upper bounds. Non-negative real numbers

Specified Value(s): 0, 0

No discards in St. Thomas/St. John were reported in SEDAR (2016). See section 2.3. A nominal amount of discard mortality is assumed in the OM.

Depletion and Discard Mortality

Sampled Parameters

Histograms of 48 simulations of depletion (spawning biomass in the last historical year over average unfished spawning biomass; D) and the fraction of discarded fish that are killed by fishing mortality (Fdisc), with vertical colored lines indicating 3 randomly drawn values.

Length-weight conversion parameters: a, b

a: Length-weight parameter alpha. Single value. Positive real number

Specified Value(s): 0

From Sagarese et al. (2018).

b: Length-weight parameter beta. Single value. Positive real number

Specified Value(s): 2.78

From Sagarese et al. (2018).

Spatial distribution and movement: Size_area_1, Frac_area_1, Prob_staying

Size_area_1: The size of area 1 relative to area 2. Uniform distribution lower and upper bounds. Positive real numbers

Specified Value(s): 0.1, 0.1

From Sagarese et al. (2018).

Frac_area_1: The fraction of the unfished biomass in stock 1. Uniform distribution lower and upper bounds. Positive real numbers

Specified Value(s): 0.1, 0.1

From Sagarese et al. (2018).

Prob_staying: The probability of inviduals in area 1 remaining in area 1 over the course of one year. Uniform distribution lower and upper bounds. Positive fraction.

Specified Value(s): 0.5, 0.6

From Sagarese et al. (2018).

Spatial & Movement

Sampled Parameters

Histograms of 48 simulations of size of area 1 (Size_area_1), fraction of unfished biomass in area 1 (Frac_area_1), and the probability of staying in area 1 in a year (Frac_area_1), with vertical colored lines indicating 3 randomly drawn values used in other plots:

Fleet Parameters

Historical years of fishing, spatial targeting: nyears, Spat_targ

nyears: The number of years for the historical spool-up simulation. Single value. Positive integer

Specified Value(s): 85

From Sagarese et al. (2018).

Spat_targ: Distribution of fishing in relation to spatial biomass: fishing distribution is proportional to B^Spat_targ. Uniform distribution lower and upper bounds. Real numbers

Specified Value(s): 1, 1.5

From Sagarese et al. (2018).

Trend in historical fishing effort (exploitation rate), interannual variability in fishing effort: EffYears, EffLower, EffUpper, Esd

EffYears: Years representing join-points (vertices) of time-varying effort. Vector. Non-negative real numbers

Adjusted effort trends for 2000-2014 for St. Thomas/St. John from SEDAR (2016), section 2.5, were used.

EffLower: Lower bound on relative effort corresponding to EffYears. Vector. Non-negative real numbers

50% of the adjusted effort (standardized to mean of 1) was used as the lower bound. Lower limit of 0 in Year 1 of the OM was assumed.

EffUpper: Upper bound on relative effort corresponding to EffYears. Vector. Non-negative real numbers

150% of the adjusted effort (standardized to mean of 1) was used as the upper bound. Upper limit of 0.3 in Year 1 of the OM was assumed.

EffYears EffLower EffUpper
1930 0.000 0.300
1936 0.466 1.400
1941 0.484 1.450
1947 0.514 1.540
1952 0.585 1.760
1958 0.617 1.850
1964 0.601 1.800
1969 0.565 1.690
1975 0.524 1.570
1980 0.595 1.780
1986 0.557 1.670
1992 0.515 1.540
1997 0.420 1.260
2003 0.382 1.150
2008 0.354 1.060
2014 0.321 0.964

Esd: Additional inter-annual variability in fishing mortality rate. Uniform distribution lower and upper bounds. Non-negative real numbers

Specified Value(s): 0.1, 0.4

From Sagarese et al. (2018).

Historical Effort

Sampled Parameters

Histograms of 48 simulations of inter-annual variability in historical fishing mortality (Esd), with vertical colored lines indicating 3 randomly drawn values used in the time-series plot:

Time-Series

Time-series plot showing 3 trends in historical fishing mortality (OM@EffUpper and OM@EffLower or OM@cpars$Find):

Annual increase in catchability, interannual variability in catchability: qinc, qcv

qinc: Average percentage change in fishing efficiency (applicable only to forward projection and input controls). Uniform distribution lower and upper bounds. Non-negative real numbers

Specified Value(s): -2, 2

From SEDAR (2016), Table 3.2.6.

qcv: Inter-annual variability in fishing efficiency (applicable only to forward projection and input controls). Uniform distribution lower and upper bounds. Non-negative real numbers

Specified Value(s): 0.1, 0.3

From SEDAR (2016), Table 3.2.6.

Future Catchability

Sampled Parameters

Histograms of 48 simulations of inter-annual variability in fishing efficiency (qcv) and average annual change in fishing efficiency (qinc), with vertical colored lines indicating 3 randomly drawn values used in the time-series plot:

Time-Series

Time-series plot showing 3 trends in future fishing efficiency (catchability):

Fishery gear length selectivity: L5, LFS, Vmaxlen, isRel

L5: Shortest length corresponding to 5 percent vulnerability. Uniform distribution lower and upper bounds. Positive real numbers

Specified Value(s): 0.96, 1.05

From Sagarese et al. (2018).

LFS: Shortest length that is fully vulnerable to fishing. Uniform distribution lower and upper bounds. Positive real numbers

Specified Value(s): 1.28, 1.4

From Sagarese et al. (2018).

Vmaxlen: The vulnerability of fish at . Uniform distribution lower and upper bounds. Fraction

Specified Value(s): 0, 0.5

From Sagarese et al. (2018).

isRel: Selectivity parameters in units of size-of-maturity (or absolute eg cm). Single value. Boolean.

Specified Value(s): TRUE

L5 and LFS are specified as fractions of length at 50% maturity.

Fishery length retention: LR5, LFR, Rmaxlen, DR

LR5: Shortest length corresponding ot 5 percent retention. Uniform distribution lower and upper bounds. Non-negative real numbers

Specified Value(s): 0, 0

Retention follows selectivity.

LFR: Shortest length that is fully retained. Uniform distribution lower and upper bounds. Non-negative real numbers

Specified Value(s): 0, 0

Retention follows selectivity.

Rmaxlen: The retention of fish at . Uniform distribution lower and upper bounds. Non-negative real numbers

Specified Value(s): 1, 1

Retention follows selectivity.

DR: Discard rate - the fraction of caught fish that are discarded. Uniform distribution lower and upper bounds. Fraction

Specified Value(s): 0, 0.1

No discards were reported in SEDAR (2016), section 2.3. A nominal amount of discarding (0-10% of total catch) is assumed for the OM.

Time-varying selectivity: SelYears, AbsSelYears, L5Lower, L5Upper, LFSLower, LFSUpper, VmaxLower, VmaxUpper

SelYears: (Optional) Years representing join-points (vertices) at which historical selectivity pattern changes. Vector. Positive real numbers

Slot not used.

AbsSelYears: (Optional) Calendar years corresponding with SelYears (eg 1951, rather than 1), used for plotting only. Vector (of same length as SelYears). Positive real numbers

Slot not used.

L5Lower: (Optional) Lower bound of L5 (use ChooseSelect function to set these). Vector. Non-negative real numbers

Slot not used.

L5Upper: (Optional) Upper bound of L5 (use ChooseSelect function to set these). Vector. Non-negative real numbers

Slot not used.

LFSLower: (Optional) Lower bound of LFS (use ChooseSelect function to set these). Vector. Non-negative real numbers

Slot not used.

LFSUpper: (Optional) Upper bound of LFS (use ChooseSelect function to set these). Vector. Non-negative real numbers

Slot not used.

VmaxLower: (Optional) Lower bound of Vmaxlen (use ChooseSelect function to set these). Vector. Fraction

Slot not used.

VmaxUpper: (Optional) Upper bound of Vmaxlen (use ChooseSelect function to set these). Vector. Fraction

Slot not used.

Current Year: CurrentYr

CurrentYr: The current calendar year (final year) of the historical simulations (eg 2011). Single value. Positive integer.

Specified Value(s): 2014

The most recent year of data for the SEDAR process.

Existing Spatial Closures: MPA

MPA: (Optional) Matrix specifying spatial closures for historical years.

Slot not used.

Obs Parameters

Overall, the observation model parameter are taken from the Imprecise_Biased model subject to a few additional changes.

Catch statistics: Cobs, Cbiascv, CAA_nsamp, CAA_ESS, CAL_nsamp, CAL_ESS

Cobs: Log-normal catch observation error expressed as a coefficient of variation. Uniform distribution lower and upper bounds. Non-negative real numbers

Specified Value(s): 0.28, 0.56

From Sagarese et al. (2018).

Cbiascv: Log-normal coefficient of variation controlling the sampling of bias in catch observations for each simulation. Uniform distribution lower and upper bounds. Non-negative real numbers

Specified Value(s): 0.28

From Sagarese et al. (2018).

CAA_nsamp: Number of catch-at-age observation per time step. Uniform distribution lower and upper bounds. Positive real numbers

Specified Value(s): 150, 200

From Sagarese et al. (2018).

CAA_ESS: Effective sample size (independent age draws) of the multinomial catch-at-age observation error model. Uniform distribution lower and upper bounds. Positive integers

Specified Value(s): 10, 20

From Sagarese et al. (2018).

CAL_nsamp: Number of catch-at-length observation per time step. Uniform distribution lower and upper bounds. Positive integers

Specified Value(s): 150, 200

From Sagarese et al. (2018).

CAL_ESS: Effective sample size (independent length draws) of the multinomial catch-at-length observation error model. Uniform distribution lower and upper bounds. Positive integers

Specified Value(s): 10, 20

From Sagarese et al. (2018).

Index imprecision, bias and hyperstability: Iobs, Ibiascv, Btobs, Btbiascv, beta

Iobs: Observation error in the relative abundance indices expressed as a coefficient of variation. Uniform distribution lower and upper bounds. Positive real numbers

Specified Value(s): 0.02, 0.03

From Sagarese et al. (2018).

Ibiascv: Not Used. Log-normal coefficient of variation controlling error in observations of relative abundance index. Uniform distribution lower and upper bounds. Positive real numbers

Specified Value(s): 0.4

This parameter is not used in this version of DLMtool.

Btobs: Log-normal coefficient of variation controlling error in observations of current stock biomass among years. Uniform distribution lower and upper bounds. Positive real numbers

Specified Value(s): 0.2, 0.5

From Sagarese et al. (2018).

Btbiascv: Uniform-log bounds for sampling persistent bias in current stock biomass. Uniform-log distribution lower and upper bounds. Positive real numbers

Specified Value(s): 0.2, 5

From Sagarese et al. (2018).

beta: A parameter controlling hyperstability/hyperdepletion where values below 1 lead to hyperstability (an index that decreases slower than true abundance) and values above 1 lead to hyperdepletion (an index that decreases more rapidly than true abundance). Uniform distribution lower and upper bounds. Positive real numbers

Specified Value(s): 1, 1

From Sagarese et al. (2018).

Bias in maturity, natural mortality rate and growth parameters: LenMbiascv, Mbiascv, Kbiascv,t0biascv, Linfbiascv

LenMbiascv: Log-normal coefficient of variation for sampling persistent bias in length at 50 percent maturity. Uniform distribution lower and upper bounds. Positive real numbers

Specified Value(s): 0.2

From Sagarese et al. (2018).

Mbiascv: Log-normal coefficient of variation for sampling persistent bias in observed natural mortality rate. Uniform distribution lower and upper bounds. Positive real numbers

Specified Value(s): 0.32

From Sagarese et al. (2018).

Kbiascv: Log-normal coefficient of variation for sampling persistent bias in observed growth parameter K. Uniform distribution lower and upper bounds. Positive real numbers

Specified Value(s): 0.35

From Sagarese et al. (2018).

t0biascv: Log-normal coefficient of variation for sampling persistent bias in observed t0. Uniform distribution lower and upper bounds. Positive real numbers

Specified Value(s): 0.5

From Sagarese et al. (2018).

Linfbiascv: Log-normal coefficient of variation for sampling persistent bias in observed maximum length. Uniform distribution lower and upper bounds. Positive real numbers

Specified Value(s): 0.12

From Sagarese et al. (2018).

Bias in length at first capture, length at full selection: LFCbiascv, LFSbiascv

LFCbiascv: Log-normal coefficient of variation for sampling persistent bias in observed length at first capture. Uniform distribution lower and upper bounds. Positive real numbers

Specified Value(s): 0.5

From Sagarese et al. (2018).

LFSbiascv: Log-normal coefficient of variation for sampling persistent bias in length-at-full selection. Uniform distribution lower and upper bounds. Positive real numbers

Specified Value(s): 0.5

From Sagarese et al. (2018).

Bias in fishery reference points, unfished biomass, FMSY, FMSY/M ratio, biomass at MSY relative to unfished: FMSYbiascv, FMSY_Mbiascv, BMSY_B0biascv

FMSYbiascv: Not used. Log-normal coefficient of variation for sampling persistent bias in FMSY. Uniform distribution lower and upper bounds. Positive real numbers

Specified Value(s): 0.2

From Sagarese et al. (2018).

FMSY_Mbiascv: Log-normal coefficient of variation for sampling persistent bias in FMSY/M. Uniform distribution lower and upper bounds. Positive real numbers

Specified Value(s): 0.11

From Sagarese et al. (2018).

BMSY_B0biascv: Log-normal coefficient of variation for sampling persistent bias in BMSY relative to unfished. Uniform distribution lower and upper bounds. Positive real numbers

Specified Value(s): 0.14

From Sagarese et al. (2018).

Management targets in terms of the index (i.e., model free), the total annual catches and absolute biomass levels: Irefbiascv, Crefbiascv, Brefbiascv

Irefbiascv: Log-normal coefficient of variation for sampling persistent bias in relative abundance index at BMSY. Uniform distribution lower and upper bounds. Positive real numbers

Specified Value(s): 0.3

From Sagarese et al. (2018).

Crefbiascv: Log-normal coefficient of variation for sampling persistent bias in MSY. Uniform distribution lower and upper bounds. Positive real numbers

Specified Value(s): 0.3

From Sagarese et al. (2018).

Brefbiascv: Log-normal coefficient of variation for sampling persistent bias in BMSY. Uniform distribution lower and upper bounds. Positive real numbers

Specified Value(s): 0.5

From Sagarese et al. (2018).

Depletion bias and imprecision: Dbiascv, Dobs

Dbiascv: Log-normal coefficient of variation for sampling persistent bias in stock depletion. Uniform distribution lower and upper bounds. Positive real numbers

Specified Value(s): 1

From Sagarese et al. (2018).

Dobs: Log-normal coefficient of variation controlling error in observations of stock depletion among years. Uniform distribution lower and upper bounds. Positive real numbers

Specified Value(s): 0.05, 0.2

From Sagarese et al. (2018).

Recruitment compensation and trend: hbiascv, Recbiascv

hbiascv: Log-normal coefficient of variation for sampling persistent bias in steepness. Uniform distribution lower and upper bounds. Positive real numbers

Specified Value(s): 0.46

From Sagarese et al. (2018).

Recbiascv: Log-normal coefficient of variation for sampling persistent bias in recent recruitment strength. Uniform distribution lower and upper bounds. Positive real numbers

Specified Value(s): 0.1, 0.3

From Sagarese et al. (2018).

Obs Plots

Observation Parameters

Catch Observations

Sampled Parameters

Histograms of 48 simulations of inter-annual variability in catch observations (Csd) and persistent bias in observed catch (Cbias), with vertical colored lines indicating 3 randomly drawn values used in other plots:

Time-Series

Depletion Observations

Sampled Parameters

Histograms of 48 simulations of inter-annual variability in depletion observations (Dobs) and persistent bias in observed depletion (Dbias), with vertical colored lines indicating 3 randomly drawn values used in other plots:

Time-Series

Abundance Observations

Sampled Parameters

Histograms of 48 simulations of inter-annual variability in abundance observations (Btobs) and persistent bias in observed abundance (Btbias), with vertical colored lines indicating 3 randomly drawn values used in other plots:

Time-Series

Index Observations

Sampled Parameters

Histograms of 48 simulations of inter-annual variability in index observations (Iobs) and hyper-stability/depletion in observed index (beta), with vertical colored lines indicating 3 randomly drawn values used in other plots:

Time-Series

Time-series plot of 3 samples of index observation error:

Plot showing an example true abundance index (blue) with 3 samples of index observation error and the hyper-stability/depletion parameter (beta):

Recruitment Observations

Sampled Parameters

Histograms of 48 simulations of inter-annual variability in index observations (Recsd) , with vertical colored lines indicating 3 randomly drawn values used in other plots:

Time-Series

Composition Observations

Sampled Parameters

Histograms of 48 simulations of catch-at-age effective sample size (CAA_ESS) and sample size (CAA_nsamp) and catch-at-length effective (CAL_ESS) and actual sample size (CAL_nsamp) with vertical colored lines indicating 3 randomly drawn values:

Parameter Observations

Sampled Parameters

Histograms of 48 simulations of bias in observed natural mortality (Mbias), von Bertalanffy growth function parameters (Linfbias, Kbias, and t0bias), length-at-maturity (lenMbias), and bias in observed length at first capture (LFCbias) and first length at full capture (LFSbias) with vertical colored lines indicating 3 randomly drawn values:

Reference Point Observations

Sampled Parameters

Histograms of 48 simulations of bias in observed FMSY/M (FMSY_Mbias), BMSY/B0 (BMSY_B0bias), reference index (Irefbias), reference abundance (Brefbias) and reference catch (Crefbias), with vertical colored lines indicating 3 randomly drawn values:

Imp Parameters

Output Control Implementation Error: TACFrac, TACSD

TACFrac: Mean fraction of TAC taken. Uniform distribution lower and upper bounds. Positive real number.

Specified Value(s): 0.75, 1.25

Inconsistent compliance and lack of enforcement has been reported (Bennett 2015), so we assume that the actual catches can be anywhere between 75-125% of the recommended TAC.

TACSD: Log-normal coefficient of variation in the fraction of Total Allowable Catch (TAC) taken. Uniform distribution lower and upper bounds. Non-negative real numbers.

Specified Value(s): 0, 0.05

We assume some inter-annual variability in following the recommended TAC.

Effort Control Implementation Error: TAEFrac, TAESD

TAEFrac: Mean fraction of TAE taken. Uniform distribution lower and upper bounds. Positive real number.

Specified Value(s): 0.75, 1.25

We have little information to inform this parameter, and set the implementation error in effort equal to the TAC implementation error.

TAESD: Log-normal coefficient of variation in the fraction of Total Allowable Effort (TAE) taken. Uniform distribution lower and upper bounds. Non-negative real numbers.

Specified Value(s): 0, 0.05

We assume some inter-annual variability in following the recommended TAE.

Size Limit Control Implementation Error: SizeLimFrac, SizeLimSD

SizeLimFrac: The real minimum size that is retained expressed as a fraction of the size. Uniform distribution lower and upper bounds. Positive real number.

Specified Value(s): 0.95, 1

We assume that, on average, some undersized fish are retained, but otherwise a size limit would be well-implemented.

SizeLimSD: Log-normal coefficient of variation controlling mismatch between a minimum size limit and the real minimum size retained. Uniform distribution lower and upper bounds. Non-negative real numbers.

Specified Value(s): 0, 0

We assume that the implementation of the size limit is consistent between years.

Imp Plots

Implementation Parameters

TAC Implementation

Sampled Parameters

Histograms of 48 simulations of inter-annual variability in TAC implementation error (TACSD) and persistent bias in TAC implementation (TACFrac), with vertical colored lines indicating 3 randomly drawn values used in other plots:

Time-Series

TAE Implementation

Sampled Parameters

Histograms of 48 simulations of inter-annual variability in TAE implementation error (TAESD) and persistent bias in TAC implementation (TAEFrac), with vertical colored lines indicating 3 randomly drawn values used in other plots:

Time-Series

Size Limit Implementation

Sampled Parameters

Histograms of 48 simulations of inter-annual variability in size limit implementation error (SizeLimSD) and persistent bias in size limit implementation (SizeLimFrac), with vertical colored lines indicating 3 randomly drawn values used in other plots:

Time-Series

Historical Simulation Plots

Historical Time-Series

Spawning Biomass

Depletion

Absolute

Vulnerable Biomass

Depletion

Absolute

Total Biomass

Depletion

Absolute

Recruitment

Relative

Absolute

Catch

Relative

Absolute

Historical Fishing Mortality

Historical Time-Series

References

Bennett, J. 2015. A summary of commercial fishing reporting compliance for Puerto Rico and the U.S. Virgin Islands for calendar years 2013 and 2014. SEDAR46-WP-06. SEDAR, North Charleston, SC. 6 pp.

Sagarese, S.R., Rios, A.B., Cass-Calay, S.L., Cummings, N.J., Bryan, M.D., Stevens, M.H., Harford, W.J., McCarthy, K.J., and Matter, V.M. 2018. Working Towards a Framework for Stock Evaluations in Data-Limited Fisheries. North American Journal of Fisheries Management 38:507-537.

Southeast Data Assessment and Review (SEDAR). 2016. Stock Assessment Report: Caribbean Data-limited Species. SEDAR, North Charleston, SC. 373 pp. Available at: http://sedarweb.org/sedar-46